During the Chipmunks' benefit concert in Paris, France, David Seville (Jason Lee) is badly injured after a falling billboard sends him flying across the stage into electric devices. After the accident, he is rushed to the hospital. Although he survives, he must stay to recuperate. He tells Alvin (voiced by Justin Long), Simon (voiced by Matthew Gray Gubler), and Theodore (voiced by Jesse McCartney) that while he is away, he has asked his Aunt Jackie to look after the Chipmunks, and that he has also made arrangements for them to go to school. They head to the airport, where the Chipmunks meet up with Jackie, who introduces her grandson Toby (Zachary Levi). After Jackie falls down a staircase and a luggage trolley crashes into her, she too is sent to a hospital, leaving Toby in charge.
Meanwhile,Ian Hawke (David Cross), former manager, now lives in the basement of Jett Records.A FedEx truck comes around the corner and a package falls out. Three singing female chipmunks, Brittany (voiced by Christina Applegate), Jeanette (voiced by Anna Faris), and Eleanor (voiced by Amy Poehler),who call themselves the Chipettes, emerge from the package and introduce themselves to Ian. Brittany says that since Ian made the Chipmunks stars, they want to be stars too. To prove their talent, they sing "Put Your Records On" and Ian hires them.
The tension between Alvin and Simon builds as Alvin hangs out with the jocks of the school, but the one who is most affected is Theodore who believes he is losing his brothers because they are fighting all the time.
The Chipmunks meet the Chipettes and gains crushes for their counterparts, but are soon forced to have a rivalry after discovering that they are with Ian. The Chipmunks and the Chipettes compete against each other in a Battle of the Bands. During lunchtime, Alvin meets up with Brittany, sharing a slightly romantic moment with her until he warns her about what Ian did to him and his brothers, but Brittany does not believe him. At the concert, the Chipettes sing "Single Ladies (Put a Ring on It)". Ian videotapes the Chipettes performance and posts it on YouTube, along with his phone number. The Chipettes win the competition and Ian became rich and famous again after Simon and Theodore forfeit the competition because Alvin failed to show up. Alvin later shows up only to see that everyone's gone. When Alvin returns home, Simon and Theodore pretend they're sleeping and ignore him. The embarrassment is too much for Theodore to take so he runs away to the zoo. When Toby and the boys find him and save him from an eagle, it brings out the best in all four of them and the Chipmunks all forgive each other and go back to "One for all, and three for one."
Soon, the Chipettes are hired as the opening act at a Britney Spears concert at the Staples Center. Unfortunately, the concert is on the same night as the school concert, so Ian decides to blow off the battle and make the Chipettes perform at the concert. However, since he focuses on Brittany the most, he wants her to perform onstage alone while Jeanette and Eleanor sing backup. Brittany refuses to sing without her sisters. So Ian threatens to barbecue them at a restaurant if they don't. At the school, Alvin gets a phone call from Brittany, saying that she and her sisters have been imprisoned in a cage. Alvin races to save them on his motorbike while Simon tells Jeanette how to open the cage on the phone. The Chipettes escape the cage, climb out of the limo, and jump onto Alvin's bike. While being chased, Brittany apologizes to Alvin for ignoring him about Ian. Ian pursues them with a remote control helicopter, but they leap off the motorbike and grab onto the helicopter. Jeanette snatches the remote and steers the helicopter away from Ian while the motorbike drives straight into Ian's groin. Simon and Theodore are about to forfeit the concert again, but Alvin and the Chipettes fly into the hall on the helicopter and sing We Are Family. Dave then comes back and watches them perform, along with Toby. At the Staples Center, Ian dresses up as Brittany wearing a wig with chipmunk ears and two sock puppets while he sings Single Ladies badly, only to be booed at the crowd and taken away by the security guards. The Chipmunks and Chipettes sing Shake your Groove Thing. The Chipmunks go back home with Dave while Toby goes back home. The Chipettes are adopted into the family and sleep at the boys' house at their request. When Daves wishes them good night and then turning the lights off, Alvin turns them back on repeatedly. After Dave trips over a skateboard, Alvin wishes him good night and turns the lights off.
The post credits show the jocks cleaning breaches with Dr. Rubin watching them.
An epilogue shows Ian being thrown into a dumpster backstage by the guards.
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